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Even if the formal transfer of sovereignty would not take place until the signature of the Treaty of Utrecht, the British Governor and garrison become the de facto rulers of the town.

During World War II (1939–1945) the Rock was again turned into a fortress and the civilian residents of Gibraltar were evacuated.

Leopards, hyenas, lynxes, wolves and bears lived among wild cattle, horses, deer, ibexes, oryxes and rhinos – all surrounded by olive trees and stone pines, with partridges and ducks overhead, tortoises in the underbrush and mussels, limpets and other shellfish in the waters.

Clive Finlayson, evolutionary biologist at the Gibraltar Museum said "this natural richness of wildlife and plants in the nearby sandy plains, woodlands, shrublands, wetlands, cliffs and coastline probably helped the Neanderthals to persist." Evidence at the cave shows the Neanderthals of Gibraltar likely used it as a shelter "for 100,000 years." Cro-Magnon man took over Gibraltar around 24,000 BCE.

for ever, without any exception or impediment whatsoever."The Treaty stipulated that no overland trade between Gibraltar and Spain was to take place, except for emergency provisions in the case that Gibraltar is unable to be supplied by sea.